Nomascus imperialis

Nomascus imperialis Temporal range: Holocene,
Illustration of the holotype specimen
Conservation status
Extinct
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Primates
Family:	Hylobatidae
Genus:	Nomascus
Species:	†N. imperialis
Binomial name
†Nomascus imperialis (Turvey et al., 2018)
Synonyms
Junzi imperialis Turvey et al, 2018

Nomascus imperialis (formerly Junzi imperialis), also known as the imperial gibbon, is a recently extinct species of gibbon that inhabited central China. It went extinct at some point after 200 BC, potentially as late as the 18th century.

The type specimen was found in an Ancient Chinese noblewoman's tomb. It is believed that when alive, during the Warring States period around 2,200 to 2,300 years ago, it was owned by Lady Xia, the mother of King Zhuangxiang of Qin and grandmother of Qin Shi Huang, the first emperor of China.

Taxonomy

The species was named based on the type specimen, an incomplete skull, in 2018 by Samuel Turvey and colleagues. It was described as Junzi imperialis, the only member of the monotypic genus Junzi, due to the distinctive morphology of its molar teeth and skull. The original generic name Junzi was coined by Turvey and his colleagues in reference to how gibbons were, in ancient China, kept by noblemen scholars, or junzi (君子), as pets. The study's authors were not allowed by Chinese authorities to sample the gibbon's DNA, and the species was thus placed in its own genus based on exclusively morphological evidence.

In 2025, a phylogenetic study of extant and extinct gibbons extracted DNA from the type specimen as well as another, older specimen, and found it to group within the genus Nomascus as the sister species to the highly endangered Hainan gibbon, one of the only extant gibbons remaining in China. It was thus moved to the genus Nomascus.

Distribution

Gibbons were depicted in historical Chinese art, here from 1427

Gibbons were recorded by ancient Chinese sources to have been found as far north as the Yellow River. Nomascus imperialis is the northernmost known gibbon, with 4,000-year-old subfossil remains of a wild individual identified from Yemao Cave in Chongqing. The northernmost record of the imperial gibbon, and of any gibbon, is of the captive individual kept by Lady Xia, which was identified from Shaanxi. Although this individual was captive, other animals in the menagerie appear to have had local origins, and it is thus assumed that this individual was caught locally and the species did naturally reach as far north as Shaanxi. This is supported by historical accounts suggesting that gibbons occurred in Shaanxi up to the 18th century.

Extinction

The imperial gibbon is the only known ape species to have gone extinct in the Holocene. The cause of its extinction remains uncertain, but likely owes to the mass deforestation occurring in central China during the Imperial period. The region inhabited by the species supported the highest human densities in the world for millennia, and extensive deforestation occurred in the range of the species by the late Imperial period. Remaining forests in the area were high-elevation, which were likely suboptimal for this species due to its potential dependency on lowland forests.

Discovery

The holotype skull was discovered when Lady Xia's tomb was opened in 2004. The living animal is thought to have been a member of Lady Xia's menagerie of luxury pets, which also included cranes, leopards, lynxes, and a black bear.